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The main purpose of the present work is to investigate the impact of soil-structure interaction on performance-based design optimization of steel moment resisting frame (MRF) structures. To this end, the seismic performance of optimally designed MRFs with rigid supports is compared with that of the optimal designs with a flexible base in the context of performance-based design. Two efficient metaheuristic algorithms, namely center of mass optimization and improved fireworks, are used to implement the optimization task. During the optimization process, nonlinear structural response-history analysis is carried out to evaluate the structural response. Two illustrative design examples of 6- and 12-story steel MRFs are presented, and it is observed that the performance-based design optimization considering soil-structure interaction decreases the structural weight and increases nonlinear structural response in comparison to rigid-based models. Therefore, in order to obtain more realistic optimal designs, soil-structure interaction should be included in the performance-based design optimization process of steel MRFs.